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Natural History
General information regarding recluse spider life history characteristics has been published.[7-9] Recluse spiders, as their names imply, are rather secretive in their habits. They are nocturnal spiders that actively attack prey and subdue it with venom. Although they don't use silk for prey capture, they do use it to line their diurnal refugia. In nature they are found under rocks and in crevices and are considered "synanthropic" meaning their populations benefit when associated with humans. When a habitat is conducive to recluses, dense populations are found. Part of the reason is that recluses are highly tolerant of conspecifics; they are one of the few spiders that can be reared communally in a jar, given that there is sufficient prey availability. As an example of their abundance, in Missouri, the author and a colleague collected 40 brown recluses in a barn within 75 minutes. In Kansas, the brown recluse is an extremely common house spider.[10] Finally, recluses have a prevalence for hiding in boxes which allows them to be transported out of their range by commerce or residential relocation. Despite this opportunity for range expansion, remarkably few verified populations have established outside the shaded area in the map shown. When they do establish, it typically is in the basement of a building and there is little expansion beyond the structure unless connected to other structures by underground pipes or passageways.
自然历史博物馆
一般资料已发表有关隐士蜘蛛生活史特征[7-9]。隐士蜘蛛，因为他们的名字所暗示的，在他们的生活习惯是如此的神秘。他们积极夜行蜘蛛毒液攻击猎物，征服它。虽然他们不使用丝绸捕获猎物，他们用它来排队日避难所。在自然界中被发现的岩石下，石缝中，被认为是“伴人”的意思时，其人口受益与人类相关。当栖息地，有利于隐士，人口密集。隐士是非常宽容的同种，其中部分原因是，他们是为数不多的社区在一个罐子里可以饲养的蜘蛛之一，有足够的猎物可用性。自己有余，作为一个例子，在密苏里州，笔者与一位同事在75分钟内收集了40棕色隐士在谷仓。在堪萨斯州，棕色隐士是一种极为常见的房子蜘蛛。[10]最后，隐士有一个藏在箱子里，使它们能够被运出其范围由商业或住宅搬迁的患病率。范围扩展，尽管有这样的机会非常少核实人口境外成立的图中的阴影区域。当他们建立，它通常是在建筑物的地下室，很少有地下管道或通道连接到其他构筑物的结构，除非有超出的现像。 http://www.ukassignment.org/uklunwen/
Despite their reclusive habits, they do occasionally bite humans. Recluses typically bite when they are trapped between flesh and another surface, as when a sleeping human rolls over on a prowling spider, or when putting on clothing or shoes containing spiders. Ways to reduce bite risk from recluse spiders include: 1) keep beds away from walls; remove bed skirts and items under the bed so that the only pathway to the bed is up the legs. 2) Keep clothing off the floor; if it is on the floor, shake it vigorously before dressing. 3) Store all intermittantly used items such as gardening clothing, baseball mitts or roller skates in spider-proof boxes or bags.
尽管他们深居简出的生活习惯，他们偶尔也会咬人。山人通常咬时，他们被困在肉体和另一面，当一个人力轧蜘蛛在窜来窜去，睡觉时，把含有蜘蛛的衣服或鞋子。隐士蜘蛛咬风险降低的方法包括：1）保持床铺远离墙壁，清除床裙，床底下和项目，让床上的唯一途径是腿。 2）保持衣着，地板，如果是在地板上，用力摇动换药前。 3）存储所有intermittantly使用的物品，如园艺服饰，棒球手套或旱冰鞋蜘蛛防盒子或袋子。
The common name "brown recluse" refers specifically to one species of spider that lives in the south central Midwest U.S. (Map). It may be found in less dense populations around the margins of the shaded area on the map. Many reports, both media and medical, forebodingly state that the brown recluse can be transported outside its range. Although this is true, it is then erroneously projected that one spider is the "tip of the iceberg" for rampant populations. In fact, verified finds of brown recluses outside of its range are rare and almost every collection is that of a single itinerant spider. Subsequent searching of the vicinity typically results in no additional recluses. The undeserved infamy that this spider has achieved outside of its range is nothing short of mind boggling. The few known instances of any recluse spider population establishing in non-native habitats typically are limited to circumscribed areas, with only rare reports of expansion from its locale.
Recluse Spider Populations The darkly shaded area of the map shows the distribution of the brown recluse spider (modified from the distribution map of Gertsch and Ennik, 1983). Additional limited populations may be found around the margins of the shaded area. The other 10 species of native recluse spiders are found in the striped area in the southwestern U.S.
The name "brown recluse" spider correctly refers only to the midwest species; additional species are known by common names such as the desert recluse, the Arizona recluse, etc. Unfortunately, non-arachnologists incorrectly lump them all under the "brown recluse" moniker. This is a potentially incorrect extrapolation because only the brown recluse has been intensively studied. All recluse species are probably capable of inflicting necrotizing bites, however, there may be behavioral and toxicological differences among the various species.
Two other spiders that have the potential to produce necrotizing wounds, though much less well-documented than the brown recluse, are the hobo spider and the yellow sac spider. The hobo spider (Tegenaria agrestis) may be found in the Pacific Northwest as far east as Montana and south into Oregon and Utah. The two yellow sac species (Cheiracanthium spp.) are found all over the United States, but probably only produce minor necrotic wounds.
Identifying the Brown Recluse Spider
One can readily learn how to identify recluse spiders with less than a minute's training. Whereas most U.S. spiders have 8 eyes, typically arranged in 2 rows of 4, the recluse spiders have 6 eyes arranged in pairs (dyads) with one anterior dyad and 2 lateral dyads (Fig. 1). All 13 species of U.S. recluses (11 native, 2 non-native) share the same eye pattern. In many publications, the violin pattern on the cephalothorax (the first body part to which the legs attach) is mentioned as a diagnostic characteristic (Fig 2). Although it is quite consistent in adult brown recluses (although it can fade in preserved specimens), many western U.S. recluse species and some young brown recluses have virtually no contrasting pigmentation in the violin region (Fig. 3, 4). In addition, recluse spiders have abdomens that are devoid of coloration pattern and their legs are covered with fine hairs but lack thickened spines.
Natural History
General information regarding recluse spider life history characteristics has been published.[7-9] Recluse spiders, as their names imply, are rather secretive in their habits. They are nocturnal spiders that actively attack prey and subdue it with venom. Although they don't use silk for prey capture, they do use it to line their diurnal refugia. In nature they are found under rocks and in crevices and are considered "synanthropic" meaning their populations benefit when associated with humans. When a habitat is conducive to recluses, dense populations are found. Part of the reason is that recluses are highly tolerant of conspecifics; they are one of the few spiders that can be reared communally in a jar, given that there is sufficient prey availability. As an example of their abundance, in Missouri, the author and a colleague collected 40 brown recluses in a barn within 75 minutes. In Kansas, the brown recluse is an extremely common house spider.[10] Finally, recluses have a prevalence for hiding in boxes which allows them to be transported out of their range by commerce or residential relocation. Despite this opportunity for range expansion, remarkably few verified populations have established outside the shaded area in the map shown. When they do establish, it typically is in the basement of a building and there is little expansion beyond the structure unless connected to other structures by underground pipes or passageways.
Despite their reclusive habits, they do occasionally bite humans. Recluses typically bite when they are trapped between flesh and another surface, as when a sleeping human rolls over on a prowling spider, or when putting on clothing or shoes containing spiders. Ways to reduce bite risk from recluse spiders include: 1) keep beds away from walls; remove bed skirts and items under the bed so that the only pathway to the bed is up the legs. 2) Keep clothing off the floor; if it is on the floor, shake it vigorously before dressing. 3) Store all intermittantly used items such as gardening clothing, baseball mitts or roller skates in spider-proof boxes or bags.
The common name "brown recluse" refers specifically to one species of spider that lives in the south central Midwest U.S. (Map). It may be found in less dense populations around the margins of the shaded area on the map. Many reports, both media and medical, forebodingly state that the brown recluse can be transported outside its range. Although this is true, it is then erroneously projected that one spider is the "tip of the iceberg" for rampant populations. In fact, verified finds of brown recluses outside of its range are rare and almost every collection is that of a single itinerant spider. Subsequent searching of the vicinity typically results in no additional recluses. The undeserved infamy that this spider has achieved outside of its range is nothing short of mind boggling. The few known instances of any recluse spider population establishing in non-native habitats typically are limited to circumscribed areas, with only rare reports of expansion from its locale.
Recluse Spider Populations The darkly shaded area of the map shows the distribution of the brown recluse spider (modified from the distribution map of Gertsch and Ennik, 1983). Additional limited populations may be found around the margins of the shaded area. The other 10 species of native recluse spiders are found in the striped area in the southwestern U.S.
The name "brown recluse" spider correctly refers only to the midwest species; additional species are known by common names such as the desert recluse, the Arizona recluse, etc. Unfortunately, non-arachnologists incorrectly lump them all under the "brown recluse" moniker. This is a potentially incorrect extrapolation because only the brown recluse has been intensively studied. All recluse species are probably capable of inflicting necrotizing bites, however, there may be behavioral and toxicological differences among the various species.
Two other spiders that have the potential to produce necrotizing wounds, though much less well-documented than the brown recluse, are the hobo spider and the yellow sac spider. The hobo spider (Tegenaria agrestis) may be found in the Pacific Northwest as far east as Montana and south into Oregon and Utah. The two yellow sac species (Cheiracanthium spp.) are found all over the United States, but probably only produce minor necrotic wounds.
Identifying the Brown Recluse Spider
One can readily learn how to identify recluse spiders with less than a minute's training. Whereas most U.S. spiders have 8 eyes, typically arranged in 2 rows of 4, the recluse spiders have 6 eyes arranged in pairs (dyads) with one anterior dyad and 2 lateral dyads (Fig. 1). All 13 species of U.S. recluses (11 native, 2 non-native) share the same eye pattern.
In many publications, the violin pattern on the cephalothorax (the first body part to which the legs attach) is mentioned as a diagnostic characteristic (Fig 2). Although it is quite consistent in adult brown recluses (although it can fade in preserved specimens), many western U.S. recluse species and some young brown recluses have virtually no contrasting pigmentation in the violin region (Fig. 3, 4). In addition, recluse spiders have abdomens that are devoid of coloration pattern and their legs are covered with fine hairs but lack thickened spines.
Other brown arachnids
Figure 11
The marbled cellar spider, Holocnemus pluchei, is repeatedly confused by the public as a brown recluse despite the fact that the brown markings are on the ventral surface of the body. (photo by R. Vetter ©)
Fearing that they might have recluse spiders, the public has brought in many other brown, eight-eyed spiders in addition to non-spider arachnids such as solpugids and daddy-long legs. The latter is differentiated from spiders in that it has one major body part as opposed to two, lacks venom glands, does not make silk and therefore, is not found in webs except as spider prey. Unfortunately, the urge to misidentify common, virtually harmless spiders as brown recluses is not restricted to the lay community.
Although bites from the brown recluse and other recluse spiders can be a source of significant morbidity, diagnoses implicating these spiders as the culprits should be restricted to those regions of the country that support populations of the spiders. On a broader scale, spider bites in general are overdiagnosed. [1] A call for more judicious evaluation has been made several times.[1-3, 11,12] Spider bites are the result of an incidental and accidental encounter between arachnid and human. In areas outside the range of recluse spiders, it has been suggested that physicians consider more strongly as differential diagnoses, many of those arthropods (fleas, hard ticks, soft ticks, mites, bedbugs, assassin bugs, etc.) that purposely seek out humans for their blood meals rather than the accidental spider encounter.[1] Wounds from these animals could stem from reactions to the animal's saliva, to toxins or to bacteria introduced while feeding. Stringent guidelines have been put forth in attempt to stem the overdiagnosis of spider bites.[1,2,12] Verified spider bites require the presence or sighting of a spider in the act of biting. In the absense of this, a necrotizing wound should be evaluated thoroughly for infectious, thrombotic, and vasculitic causes. Without verification, the diagnosis of necrotizing spider bite should be one of exclusion.
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